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Astanga Hrudaya

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शुक्राश्मरी तु महतां जायते शुक्रधारणात् । स्थानाच्च्युतममुक्तं हि मुष्कयोरन्तरेऽनिलः ॥ १६ ॥

Seminal stone (Sukrasmari) gets formed in adults due to the suppression of the flow of semen. When the semen has left its seat (of production) but is not ejaculated, the Anila (Vata) Dosha staying inside the scrotum gets aggravated.

english translation