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Charak Samhita

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मातापित्रोर्बीजदोषादशुभैश्चाकृतात्मनः । गर्भस्थस्य यदा दोषाः प्राप्य रेतोवहाःसिराः ॥१८९॥

Dosha due to beejadosha (genetic defects) of parents and ominous past conduct when penetrate into the virile semen system,

english translation

Chapter 30

Verse 190